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The occipital bone () is a and the main of the occiput (back and lower part of the ). It is in shape and curved on itself like a shallow dish. The occipital bone lies over the of the . At the base of the skull in the occipital bone, there is a large oval opening called the , which allows the passage of the .

Like the other cranial bones, it is classed as a . Due to its many attachments and features, the occipital bone is described in terms of separate parts. From its front to the back is the basilar part, also called the basioccipital, at the sides of the foramen magnum are the lateral parts, also called the exoccipitals, and the back is named as the squamous part. The basilar part is a thick, somewhat quadrilateral piece in front of the foramen magnum and directed towards the . The squamous part is the curved, expanded plate behind the foramen magnum and is the largest part of the occipital bone.

Due to its embryonic derivation from paraxial mesoderm (as opposed to neural crest, from which many other craniofacial bones are derived), it has been posited that "the occipital bone as a whole could be considered as a giant vertebra enlarged to support the brain."


Structure
The occipital bone, like the other seven , has outer and inner layers (also called plates or tables) of cortical bone tissue between which is the cancellous bone tissue known in the cranial bones as diploë. The bone is especially thick at the ridges, protuberances, condyles, and anterior part of the basilar part; in the inferior cerebellar fossae it is thin, semitransparent, and without diploë.


Outer surface
Near the middle of the outer surface of the squamous part of the occipital (the largest part) there is a prominence – the external occipital protuberance. The highest point of this is called the inion.

From the inion, along the midline of the squamous part until the foramen magnum, runs a ridge – the external occipital crest (also called the medial nuchal line) and this gives attachment to the .

Running across the outside of the occipital bone are three curved lines and one line (the medial line) that runs down to the foramen magnum. These are known as the which give attachment to various ligaments and muscles. They are named as the highest, superior and inferior nuchal lines. The inferior nuchal line runs across the midpoint of the median nuchal line. The area above the highest nuchal line is termed the and the area below this line is termed the .



Inner surface
The inner surface of the occipital bone forms the base of the posterior cranial fossa. The is a large hole situated in the middle, with the clivus, a smooth part of the occipital bone travelling upwards in front of it. The median internal occipital crest travels behind it to the internal occipital protuberance, and serves as a point of attachment to the .

To the sides of the sitting at the junction between the lateral and base of the occipital bone are the hypoglossal canals. Further out, at each junction between the occipital and petrous portion of the temporal bone lies a .

The inner surface of the occipital bone is marked by dividing lines as shallow ridges, that form four fossae or depressions. The lines are called the cruciform (cross-shaped) eminence.

At the midpoint where the lines intersect a raised part is formed called the internal occipital protuberance. From each side of this eminence runs a groove for the transverse sinuses.

There are two midline skull landmarks at the . The basion is the most anterior point of the opening and the opisthion is the point on the opposite posterior part. The basion lines up with the dens.


Foramen magnum
The foramen magnum () is a large oval longest front to back; it is wider behind than in front where it is encroached upon by the occipital condyles. The clivus, a smooth bony section, travels upwards on the front surface of the foramen, and the median internal occipital crest travels behind it.

Through the foramen passes the medulla oblongata and its membranes, the , the , the anterior and posterior spinal arteries, the tectorial membrane and the .


Angles
The superior angle of the occipital bone articulates with the occipital angles of the and, in the fetal skull, corresponds in position with the posterior fontanelle.

The lateral angles are situated at the extremities of the groove for the transverse sinuses: each is received into the interval between the mastoid angle of the parietal bone, and the mastoid portion of the temporal bone.

The inferior angle is fused with the body of the sphenoid bone.


Borders
The superior borders extend from the superior to the lateral angles: they are deeply serrated for articulation with the occipital borders of the parietals, and form by this union the lambdoidal suture.

The inferior borders extend from the lateral angles to the inferior angle; the upper half of each articulates with the mastoid portion of the corresponding , the lower half with the petrous part of the same bone.

These two portions of the inferior border are separated from one another by the , the notch on the anterior surface of which forms the posterior part of the .

===Sutures===

The joins the occipital bone to the .

The occipitomastoid suture joins the occipital bone and of the .

The sphenobasilar suture joins the basilar part of the occipital bone and the back of the body.

The petrous-basilar suture joins the side edge of the basilar part of the occipital bone to the of the .


Development
The Fig. of the squamous part of the occipital bone is developed in membrane, and may remain separate throughout life when it constitutes the interparietal bone; the rest of the bone is developed in cartilage.

The number of nuclei for the occipital plane is usually given as four, two appearing near the middle line about the second month, and two some little distance from the middle line about the third month of fetal life.

The of the squamous part is ossified from two centers, which appear about the seventh week of fetal life and soon unite to form a single piece.

Union of the upper and lower portions of the squamous part takes place in the third month of fetal life.

An occasional centre (Kerckring) appears in the posterior margin of the foramen magnum during the fifth month; this forms a separate ossicle (sometimes double) which unites with the rest of the squamous part before birth.

Each of the lateral parts begins to from a single center during the eighth week of fetal life. The basilar portion is ossified from two centers, one in front of the other; these appear about the sixth week of fetal life and rapidly coalesce.

The occipital plane is said to be ossified from two centers and the basilar portion from one.

About the fourth year the squamous part and the two lateral parts unite, and by about the sixth year the bone consists of a single piece. Between the 18th and 25th years the occipital and become united, forming a single bone.


Clinical significance
Trauma to the occiput can cause a fracture of the base of the skull, called a basilar skull fracture. The basion-dens line as seen on a is the distance between the basion and the top of the dens, used in the diagnosis of dissociation injuries.

can cause a prominent occiput as found in , and Beckwith–Wiedemann syndrome.

The identification of the location of the occiput is important in .


Etymology
Occipital stems from Latin "back of the skull", from ob "against, behind" + caput "head". Distinguished from (anterior part of the skull)."occipital" A Dictionary of Zoology. Ed. Michael Allaby. Oxford University Press 2009


Other animals
In many animals these parts stay separate throughout life; for example, in the dog as four parts: squamous part (supraoccipital); lateral parts–left and right parts (exoccipital); basilar part (basioccipital).

The occipital bone is part of the , the most basal portion of the skull. In and , the occipital does not form as a separate element, but remains part of the throughout life. In most higher vertebrates, the foramen magnum is surrounded by a ring of four bones.

The basioccipital lies in front of the opening, the two exoccipital condyles lie to either side, and the larger supraoccipital lies to the posterior, and forms at least part of the rear of the cranium. In many and , the supraoccipital is never ossified, and remains as throughout life. In primitive forms the basioccipital and exoccipitals somewhat resemble the centrum and neural arches of a , and form in a similar manner in the embryo. Together, these latter bones usually form a single concave circular for the articulation of the first .

(1977). 003910284X, Holt-Saunders International. 003910284X

In , however, the condyle has divided in two, a pattern otherwise seen only in a few amphibians.

Most mammals also have a single fused occipital bone, formed from the four separate elements around the foramen magnum, along with the paired bones that form the rear of the in other vertebrates.

== Additional images ==

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